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"In mice, athorny pyramidal cells were also visualized using Golgi staining or viral injection, where they were found to be negative for the vesicular GABA transporter (VGAT), a ubiquitous inhibitory marker, and positive for the glutamatergic markers VGLUT1 and VGLUT2, indicating that they are not GABAergic (Supplementary Figs. 2-4)."
"Supplementary Figure 4
Athorny neurons are positive for glutamatergic markers VGLUT1 & VGLUT2
... c) Single optical sections through the soma of the cells shown above in (b) where the GFP positive somatic region is also positive for the glutamatergic markers VGLUT1 and VGLUT2 (n = 3 cells)."